- These are also giant chromosomes but relatively smaller than lampbrush chromosomes, found in the larvae of certain dipterans. Such banded chromosomes occur in the larval salivary glands, midgut epithelium, and rectum and Malpighian tubules of various genera (Drosophila, Sciara, Rhynchosciara, and Chironomus). In these larvae the salivary glands contain salivary cells so large in size that they can easily be seen with the lens power of a dissecting microscope.Nuclei of these cells are much larger than those of ordinary cells being generally about 25µ in diameter, and chromosomes in nuclei are so large that they are 50 to 200 times as large as chromosomes in other body cells of the organism.
- They were first observed in 1881 by E.G. Balbiani in Chironomus and were studied by Korschelt (1884) and Corney (1884). Heitz and Bauer in 1933 studied these giant chromosomes in Bibio hortulanus larvae, while Painter (1933) described them in salivary glands of Drosophila.

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